(g) Phase shift showing absolute lag distance corresponding to in-phase peak in Venus::HES5 vs NGN2::mScarlet-I cross-correlation function of spatial intensity profiles from the same slice. 34 individual data points from 6 slices, 2 experiments; red line indicates average inter-nuclear distance in D-V; bars show mean±SD; 2-tailed Mann-Whitney test not significant, p=0.32. 