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BiologyOccur in sandy and muddy, sometimes rocky, pools of headwaters, creeks, and small to medium rivers; and in sandy shores of lakes (Ref. 5723, 10294); also found in streams (Ref. 10294). Adults feed on midge larvae, mayfly nymphs, caddis larvae, and microcrustaceans; young on entomostracans and tiny midge larvae (Ref. 10294). Eggs are found clustered on underside of stone and guarded by males (Ref. 7043).