  The largest galaxies, and in particular central galaxies in clusters, offer
unique insight into understanding the mechanism for the growth of nuclear black
holes. We present Hubble Space Telescope kinematics for NGC1399, the central
galaxy in Fornax. We find the best-fit model contains a black hole of 5.1 +-0.7
x 10^8 Msun (at a distance of 21.1 Mpc), a factor of over 2 below the
correlation of black hole mass and velocity dispersion. We also find a dramatic
signature for central tangential anisotropy. The velocity profiles on adjacent
sides 0.5" away from the nucleus show strong bimodality, and the central
spectrum shows a large drop in the dispersion. Both of these observations point
to an orbital distribution that is tangentially biased. The best-fit orbital
model suggests a ratio of the tangential to radial internal velocity
dispersions of three. This ratio is the largest seen in any galaxy to date and
will provide an important measure for the mode by which the central black hole
has grown.
