  We discuss multiplicity fluctuations of charged particles produced in nuclear
collisions measured event-by-event by the NA49 experiment at CERN SPS within
the Glauber Monte Carlo approach. We use the concepts of wounded nucleons and
wounded quarks in the mechanism of multiparticle production to characterize
multiplicity fluctuations expressed by the scaled variance of multiplicity
distribution. Although Wounded Nucleon Model correctly reproduce the centrality
dependence of the average multiplicity in Pb+Pb collisions, it completely fails
in description of corresponding centrality dependence of scaled variance of
multiplicity distribution. Using subnucleonic degrees of freedom, i.e. wounded
quarks within Wounded Quark Model, it is possible to describe quite well the
multiplicity distribution of charged particles produced in proton+proton
interactions. However, the Wounded Quark Model with parameters describing
multiplicity distribution of particles produced in proton+proton interactions
substantially exceeds the average multiplicity of charged particles produced in
Pb+Pb collisions. To obtain values of average multiplicities close to those
experimentally measured in Pb+Pb collisions, the concept of shadowed quark
sources is implemented. Wounded Quark Model with implemented shadowing source
scenario reproduces the centrality dependence of scaled variance of
multiplicity distribution of charged particles produced in Pb+Pb collisions in
the range from the most central to mid-peripheral interactions.
