  We discuss a diffusively perturbed predator-prey system. Freedman and
Wolkowicz showed that the corresponding ODE can have a periodic solution that
bifurcates from a homoclinic loop. When the diffusion coefficients are large,
this solution represents a stable, spatially homogeneous time-periodic solution
of the PDE. We show that when the diffusion coefficients become small, the
spatially homogeneous periodic solution becomes unstable and bifurcates into
spatially nonhomogeneous periodic solutions.
  The nature of the bifurcation is determined by the twistedness of an
equilibrium/homoclinic bifurcation that occurs as the diffusion coefficients
decrease. In the nontwisted case two spatially nonhomogeneous simple periodic
solutions of equal period are generated, while in the twisted case a unique
spatially nonhomogeneous double periodic solution is generated through
period-doubling.
  Key Words: Reaction-diffusion equations; predator-prey systems; homoclinic
bifurcations; periodic solutions.
