  Let $\pi:M\to B$ be a proper holomorphic submersion between complex manifolds
and ${\cal E}$ a holomorphic bundle on $M$. We study and describe explicitly
the torsion subsheaf $\mathrm{Tors}(R^1\pi_*({\cal E}))$ of the first direct
image $R^1\pi_*(\mathcal{E})$ under the assumption $R^0\pi_*(\mathcal{E})=0$.
We give two applications of our results. The first concerns the locus of points
in the base of a generically versal family of complex surfaces where the family
is non-versal. The second application is a vanishing result for
$H^0(\mathrm{Tors}(R^1\pi_*(\mathcal{E})))$ in a concrete situation related to
our program to prove the existence of curves on class VII surfaces.
