  Planetary obliquity and eccentricity influence climate by shaping the spatial
and temporal patterns of stellar energy incident at a planet's surface,
affecting both the annual mean climate and magnitude of seasonal variability.
Previous work has demonstrated the importance of both planetary obliquity and
eccentricity for climate and habitability, but most studies have not explicitly
modeled the response of life to these parameters. While exaggerated seasons may
be stressful to some types of life, a recent study found an increase in marine
biological activity for moderately high obliquities <45$^{\circ}$ assuming an
Earth-like eccentricity. However, it is unclear how life might respond to
obliquities >45$^{\circ}$, eccentricities much larger than Earth's, or the
combination of both. To address this gap, we use cGENIE-PlaSim, a 3-D marine
biogeochemical model coupled to an atmospheric general circulation model, to
investigate the response of Earth-like marine life to a large range of
obliquities (0-90$^{\circ}$) and eccentricities (0-0.4). We find that marine
biological activity increases with both increasing obliquity and eccentricity
across the parameter space we considered, including the combination of high
obliquity and high eccentricity. We discuss these results in the context of
remote biosignatures, and we argue that planets with high obliquity and/or
eccentricity may be superhabitable worlds that are particularly favorable for
exoplanet life detection.
