  Real social interactions occur on networks in which each individual is
connected to some, but not all, of others. In social dilemma games with a fixed
population size, heterogeneity in the number of contacts per player is known to
promote evolution of cooperation. Under a common assumption of positively
biased payoff structure, well-connected players earn much by playing
frequently, and cooperation once adopted by well-connected players is
unbeatable and spreads to others. However, maintaining a social contact can be
costly, which would prevent local payoffs from being positively biased. In
replicator-type evolutionary dynamics, it is shown that even a relatively small
participation cost extinguishes the merit of heterogeneous networks in terms of
cooperation. In this situation, more connected players earn less so that they
are no longer spreaders of cooperation. Instead, those with fewer contacts win
and guide the evolution. The participation cost, or the baseline payoff, is
irrelevant in homogeneous populations but is essential for evolutionary games
on heterogeneous networks.
