  We study the consequences of a homogeneous dark energy fluid having a
non-vanishing velocity with respect to the matter and radiation large-scale
rest frames. We consider homogeneous anisotropic cosmological models with four
fluids (baryons, radiation, dark matter and dark energy) whose velocities can
differ from each other. Performing a perturbative calculation up to second
order in the velocities, we obtain the contribution of the anisotropies
generated by the fluids motion to the CMB quadrupole and compare with
observations. We also consider the exact problem for arbitrary velocities and
solve the corresponding equations numerically for different dark energy models.
We find that models whose equation of state is initially stiffer than
radiation, as for instance some tracking models, are unstable against velocity
perturbations, thus spoiling the late-time predictions for the energy
densities. In the case of scaling models, the contributions to the quadrupole
can be non-negligible for a wide range of initial conditions. We also consider
fluids moving at the speed of light (null fluids) with positive energy and show
that, without assuming any particular equation of state, they generically act
as a cosmological constant at late times. We find the parameter region for
which the models considered could be compatible with the measured (low)
quadrupole.
