  We present an investigation of new optical and ultraviolet spectra of the
mass donor star in the massive X-ray binary LS 5039. The optical band spectral
line strengths indicate that the atmosphere is N-rich and C-poor, and we
classify the stellar spectrum as type ON6.5 V((f)). The N-strong and C-weak
pattern is also found in the stellar wind P Cygni lines of N V 1240 and C IV
1550. We suggest that the N-enrichment may result from internal mixing if the
O-star was born as a rapid rotator, or the O-star may have accreted N-rich gas
prior to a common-envelope interaction with the progenitor of the supernova. We
re-evaluated the orbital elements to find an orbital period of P=4.4267 +/-
0.0010 d. We compared the spectral line profiles with new non-LTE,
line-blanketed model spectra, from which we derive an effective temperature
T_eff = 37.5 +/- 1.7 kK, gravity log g = 4.0 +/- 0.1, and projected rotational
velocity V sin i = 140 +/- 8 km/s. We fit the UV, optical, and IR flux
distribution using a model spectrum and extinction law with parameters E(B-V)=
1.28 +/- 0.02 and R= 3.18 +/- 0.07. We confirm the co-variability of the
observed X-ray flux and stellar wind mass loss rate derived from the H-alpha
profile, which supports the wind accretion scenario for the X-ray production in
LS 5039. Wind accretion models indicate that the compact companion has a mass
M_X/M_sun = 1.4 +/- 0.4, consistent with its identification as a neutron star.
The observed eccentricity and runaway velocity of the binary can only be
reconciled if the neutron star received a modest kick velocity due to a slight
asymmetry in the supernova explosion (during which >5 solar masses was
ejected).
