  Cooperation is ubiquitous in biological and social systems. Previous studies
revealed that a preference toward similar appearance promotes cooperation, a
phenomenon called tag-mediated cooperation or communitarian cooperation. This
effect is enhanced when a spatial structure is incorporated, because space
allows agents sharing an identical tag to regroup to form locally cooperative
clusters. In spatially distributed settings, one can also consider migration of
organisms, which has a potential to further promote evolution of cooperation by
facilitating spatial clustering. However, it has not yet been considered in
spatial tag-mediated cooperation models. Here we show, using computer
simulations of a spatial model of evolutionary games with organismal migration,
that tag-based segregation and homophilic cooperation arise for a wide range of
parameters. In the meantime, our results also show another evolutionarily
stable outcome, where a high level of heterophilic cooperation is maintained in
spatially well-mixed patterns. We found that these two different forms of
tag-mediated cooperation appear alternately as the parameter for temptation to
defect is increased.
