  The physical origins of the Fundamental Plane (FP) scaling relations are
investigated for early-type galaxies observed at optical and near-infrared
wavelengths. The slope for the FP is shown to increase systematically with
wavelength from the U-band through the K-band. A distance-independent
construction of the observables is described which provides an accurate
measurement of the change in the FP slope between any pair of bandpasses. The
variation of the FP slope with wavelength is strong evidence of systematic
variations in stellar content along the elliptical galaxy sequence. The
intercept of the diagnostic relationship between log(D_K/D_V) and log(sigma_0)
shows no significant dependence on environment within the uncertainties of the
Galactic extinction corrections, demonstrating the universality of the stellar
populations contributions at the level of Delta(V-K)=0.03 mag to the zero-point
of the global scaling relations.
  Several other constraints on the properties of early-type galaxies --- the
slope of the Mg_2-sigma_0 relation, the effects of stellar populations
gradients, and deviations of early-type galaxies from a dynamically homologous
family --- are included to construct an empirical, self-consistent model which
provides a complete picture of the underlying physical properties which are
varying along the early-type galaxy sequence. This empirical approach
demonstrates that there are significant systematic variations in both age and
metallicity along the elliptical galaxy sequence, and that a small, but
systematic, breaking of dynamical homology (or a similar, wavelength
independent effect) is required. Predictions for the evolution of the slope of
the FP with redshift are described. [abriged]
