  To understand the relationship between the chirality of individual cells and
that of tissues and organisms, we have developed a chiral polarized particle
model to investigate the movement of cell populations on substrates. Our model
analysis indicates that cells with the same chirality can form distinct chiral
patterns on ring-shaped or rectangular substrates. Our model analysis also
reveals the importance of coordination between boundary features and individual
cellular chirality in regulating the movement of cell populations. This work
provides valuable insights into comprehending the intricate connection between
the chirality of single cells and that of tissues and organisms.
