  The fraction of stellar mass contained in globular clusters (GCs), also
measured by number as the specific frequency, is a fundamental quantity that
reflects both a galaxy's early star formation and its entire merging history.
We present specific frequencies, luminosities, and mass fractions for the
globular cluster systems of 100 early-type galaxies in the ACS Virgo Cluster
Survey, the largest homogeneous catalog of its kind. We find that 1) GC mass
fractions can be high in both giants and dwarfs, but are universally low in
galaxies with intermediate luminosities. 2) The behavior of specific frequency
across galaxy mass is dominated by the blue GCs. 3) The GC fractions of
low-mass galaxies exhibit a dependence on environment. Nearly all dwarf
galaxies with high GC fractions are within 1 Mpc of the cD galaxy M87,
presenting the first strong evidence that GC formation in dwarfs is biased
toward dense environments. 4) GC formation in central dwarfs is biased because
their stars form earliest and most intensely. Comparisons to the Millennium
Simulation show that central dwarfs have older stellar populations and form
more stars at higher star formation rates (SFRs) and SFR surface densities. The
SFR surface density in simulated dwarfs peaks before the total SFR, naturally
producing GC populations that are older and more metal-poor than the field
stars. 5) Dwarfs within ~40 kpc of the giant ellipticals M87 and M49 are red
and have few or no GCs, suggesting that they have been tidally stripped and
have contributed their GCs to the halos of their giant neighbors. The central
dwarfs with high GC mass fractions are thus likely to be the survivors most
similar to the protogalaxies that assembled the rich M87 globular cluster
system.(Abridged)
